New and little-known genera and species of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) from New Guinea.

Two formerly monotypic subgenera of Scytocera Redtenbacher, 1891, namely Morocera Gorochov, 2016 stat. nov. and Papucera Gorochov, 2016 stat. nov. are raised to full genus status. The first male of Morocera loriae (Griffini, 1908) and two new species of this genus are described: M. nigrifrons sp. nov. and M. peniculosa sp. nov. In the genus Papucera, formerly only known from P. smaragdifrons Naskrecki & Rentz (2010), P. dentata sp. nov. is described as new species. Four new genera are proposed to accommodate species with unique characters, namely Cuspicera gen. nov. for two species, C. luxoria spec. nov. (typus generis) and C. alata sp. nov., Kaindiagraecia gen. nov. for K. bilobata sp. nov., Glennagraecia gen. nov. for G. curvata sp. nov. and Angustocera gen. nov. for A. plana sp. nov. All data are based on specimens found in historical museum collections in Europe and North America.

Revision of the genus Philmontis Willemse, 1966 and description of a new genus Philmontoides gen. nov. from New Guinea (Orthoptera: Tettigoniidae; Conocephalinae).

The genera Philmontis Willemse, 1966 and Philmontoides gen. nov. are revised. Both genera are of similar general appearance and share a similar color pattern. They differ in the length of the male pronotum that covers the stridulatory area in Philmontis but leaves it free in Philmontoides, by the length of the prosternal spines, by the absence or presence of spines on the inner margin of the hind femur and the hind knees being provided with a single or with two apical spines. The male titillators of Philmontis are often elongate and rather simple, but always terminate into a small sclerotized disc that is missing in Philmontoides and in which the titillators of most species show striking modifications of the apical areas differing among species. The ovipositor is regularly curved to an acute end in Philmontis but has the dorsal valves expanded around mid-length in Philmontoides. The affinity of Nicsara forcipata Willemse, 1966 with the type species of Philmontis (P. nigrofasciatus Willemse, 1966) is confirmed on basis of the homology of the male cerci and titillators in both species. P. minimus Willemse, 1966, originally described as forma of P. nigrofasciatus, is given full species status. Eleven species are described as new in the genus Philmontis Willemse, 1966: Philmontis angustus sp. nov., P. murmur sp. nov., P. banz sp. nov., P. flexus sp. nov., P. extensus sp. nov., P. spinosus sp. nov., P. angulatus sp. nov., P. profusus sp. nov., P. robustus sp. nov., P. pandus sp. nov. and P. pumilus sp. nov.Lobaspis hageni Dohrn, 1905 is selected as type species of the genus Philmontoides gen. nov. Two species, Nicsara affinis Willemse, 1966 and Philmontis lobatus Naskrecki Rentz, 2010, are newly combined with Philmontoides. Six species are described as new in that genus: Philmontoides globosus sp. nov., P. disjunctus sp. nov., P. striatus sp. nov., P. geminus sp. nov., P. commodus sp. nov., P. wau sp. nov.

Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini).

The genus Habetia Kirby, 1906 previously known from only four species or subspecies is revised based on unidentified specimens found in museum collections. H. multispinulosa Griffini, 1908 is given full species status, while H. pictifrons Karny, 1911 is removed from Habetia based on male and female characters and becomes the type species of the new genus Parahabetia gen. nov. Sixteen new species are described in Habetia, and one new species and two new subspecies in Parahabetia. The most significant characters for the differentiation of species of Habetia are male tenth abdominal tergite, cerci, subgenital plate, and male internal genitalia with the titillators that basically form a pair of roughly U-shaped sclerites that vary between species from simple, curved sclerites to modified structures with additional sclerotized branches and the sclerites can be connected with septa to form complex shapes. In contrast, males of Parahabetia have elongate titillators provided at end with a crest. In the larger species of Habetia the stridulatory file on the underside of the male left tegmen has a step in pre-apical area that is missing in the smaller species; one species with a weak step is intermediate between both groups. Unique modifications of single species in Habetia are an apical dorsal extension of the male subgenital plate in H. imitatrix Karny, 1912 and a hardened phallus sheath between phallus and subgenital plate in H. tuta sp. nov. Females of Habetia have straight ovipositors that vary in length and in the shape of the dorsal margin between species. Females can also be differentiated by species specific modifications of the subgenital plate. In Parahabetia the ovipositor is rather short and curved similar to the condition in the genus Philmontis Willemse, 1966 but has the apical area slightly recurved. Keys to the species of both genera are provided. Habetia species have been found throughout New Guinea except so far for Papua Barat; although they were mainly found in the North and the East. Parahabetia species are currently only known from the East of New Guinea. The following species and subspecies are described as new: Habetia bivittata sp. nov., H. curvata sp. nov., H. dentata sp. nov., H. elevata sp. nov., H. kondiu sp. nov., H. lalibu sp. nov., H. nigricauda sp. nov., H. pallida sp. nov., H. pedala sp. nov., H. pilleata sp. nov., H. pinnigera sp. nov., H. quatrispina sp. nov., H. simbai sp. nov., H. sororcula sp. nov., H. tuta sp. nov., H. wau sp. nov., Parahabetia bispinosa sp. nov., P. pictifrons obtusa ssp. nov., P. pictifrons acuta ssp. nov.

Bioacoustics and systematics of Mecopoda (and related forms) from South East Asia and adjacent areas (Orthoptera, Tettigonioidea, Mecopodinae) including some chromosome data.

Bush-crickets (or katydids) of the genus Mecopoda are relatively large insects well-known for their sounds for centuries. Bioacoustic studies in India and China revealed a surprisingly large diversity of sound patterns. We extend these studies into the tropics of South East Asia using integrative taxonomy, combining song analysis, morphology of sound producing organs and male genitalia as well as chromosomes, to get a better understanding of the phylogeny and evolution of this widespread group. Besides the closely related genus Eumecopoda, the genus Mecopoda contains some isolated species and a large group of species which we assign to the Mecopoda elongata group. Some species of this group have broad tegmina and stridulatory files with different tooth spacing patterns and produce continuous, often relatively complicated, trill-like songs. The species of another subgroup with narrower wings have all similar files. Their songs consist of echemes (groups of syllables) which differ in syllable number and syllable repetition rate and also in echeme repetition rate. Our results show that South East Asia harbours a large and certainly not yet fully explored number of Mecopoda species which are most easily and clearly identified by song. Based on the data, five new forms are described: Mecopoda mahindai Heller sp. nov., Mecopoda paucidens Ingrisch, Su Heller sp. nov., Mecopoda sismondoi Heller sp. nov., Mecopoda niponensis vietnamica Heller Korsunovskaya subsp. nov., Eumecopoda cyrtoscelis zhantievi Heller subsp. nov. In addition, some taxonomic changes are proposed: Eumecopoda Hebard, 1922 stat. rev., Paramecopoda Gorochov, 2020, syn. nov. of Eumecopoda Hebard, 1922, Mecopoda javana (Johansson, 1763) stat. nov. (neotype selected) with M. javana minahasa Gorochov, 2020 stat. nov., M. javana darevskyi Gorochov, 2020 stat. nov., M. javana buru Gorochov, 2020 stat. nov., Mecopoda macassariensis (Haan, 1843) stat. rev., Mecopoda ampla malayensis Gorochov, 2020 syn. nov., Mecopada ampla javaensis Gorochov, 2020 syn. nov., Mecopoda fallax aequatorialis Gorochov, 2020 syn. nov., the last three are all synonyms of Mecopoda himalaya Liu, 2020, Mecopoda yunnana Liu 2020, stat. nov.

A new species of Heminicsara (Orthoptera: Tettigoniidae: Agraeciini) from West Papua (New Guinea).

Heminicsara Karny, 1912 is a katydid genus of Agraeciini from the Axylus genus group. It currently comprises 62 species from mainly New Guinea and surrounding archipelagos. Based on recent fieldwork in Lobo in West Papua, Indonesia, a new species of Heminicsara is described here: Heminicsara incrassata sp. nov. It is most readily characterised from congeners and other species of the Axylus genus group by the male tenth abdominal tergite forming a large shield-shaped plate. This represents the first species of Heminicsara described and known from the south-west of New Guinea.

New species and genera of Agraeciini (Orthoptera, Tettigoniidae, Conocephalinae) from New Guinea, Solomon Islands and Sulawesi.

The formerly monotypic genera Tamolana Kuthy, 1910 and Ingrischia Naskrecki Rentz, 2010 are revised. Nanoagraecia gen. nov. is introduced as new genus, and Strictonicsara subgen. nov. as new subgenus of Pseudonicsara Karny, 1912. Five new species of Salomona Blanchard, 1853, one new species of Paramacroxiphus Willemse, 1961, and one new species of Jambiliara Ingrisch, 1998 are described. An updated key to the species of Paramacroxiphus is given. The following species are described as new: Tamolana extensa sp. nov., Tamolana arborea sp. nov., Tamolana malu sp. nov., Ingrischia acuticeps sp. nov., Ingrischia brevicona sp. nov., Ingrischia carinata sp. nov., Ingrischia signifer sp. nov., Ingrischia samberi sp. nov., Salomona brevivertex sp. nov., Salomona gumunang sp. nov., Salomona fuscifrons sp. nov., Salomona longicornis sp. nov., Salomona lanigera sp. nov., Paramacroxiphus bistylatus sp. nov., Pseudonicsara (Strictonicsara) brachyptera sp. nov., Pseudonicsara (Strictonicsara) rugosa sp. nov., Nanoagraecia gibberosa sp. nov., Jambiliara rotunda sp. nov. A subdivision of the female subgenital plate of Salomona into two sclerites plus a pair of lateral appendages is described and compared to a similar modification in the genus Ingrischia.

New species of Ornebius (Orthoptera: Mogoplistidae; Mogoplistinae) from Siargao Island of the Philippines.

Two new species of Ornebius are described from Siargao Island, Mindanao of the Philippines: O. alvarezi sp. nov. and O. bioculatus sp. nov. These sympatric scaly crickets have diverging genitalia morphology, with O. alvarezi sp. nov. resembling congeners from Borneo whereas O. bioculatus sp. nov. resembling congeners from Sulawesi. With the new findings, the number of Ornebius species known from the Philippines increases to four.

Revision of the genus Rhytidaspis Redtenbacher, 1891 including the description of a new genus Haudrhytidaspis gen. nov. (Orthoptera: Tettigoniidae: Conocephalinae).

The genus Rhytidaspis Redtenbacher, 1891 is revised. The species R. fusca (Karny, 1911) is removed from the genus and combined with Haudrhytidaspis gen. nov. as type species, leaving only the type species R. picta Redtenbacher, 1891 in Rhytidaspis. The shapes of the male cerci and the complex structure of the male titillators in Rhytidaspis allows to differentiate between seven species and two subspecies of which six species and two subspecies are new to science: R. arfak sp. n., R. camela sp. n., R. genyem sp. n., R. nigropunctata sp. n., R. ornata sp. n., R. variata sp. n., R. variata brevis ssp. n., R. variata meja ssp. n.. A dichotomic key to species and a distribution map are provided. All records of Rhytidaspis known so far are from western New Guinea: Papua and Papua Barat (Indonesia) including Waigeo Island. Haudrhytidaspis fusca (Karny, 1911) had been described from New Guinea without precision.

New taxa and records of Gryllacrididae (Orthoptera, Stenopelmatoidea) from South East Asia and New Guinea with a key to the genera.

New records of Gryllacrididae from South East Asia to New Guinea are reported, new taxa are described and type specimens from European collections redescribed. The wing venation of the tegmina and its variation within the family is discussed. A new key to the 50 genera and subgenera of Gryllacrididae occurring in the area from South East Asia to New Guinea is provided. Six genera and two subgenera are introduced as new: Angustogryllacris gen. nov., Minigryllacris gen. nov., Monseremus gen. nov., Siamgryllacris gen. nov., Paraneanias gen. nov., Plexigryllacris gen. nov., Pseudasarca subgen. nov. of Asarcogryllacris Karny, 1937, Xiphilarnaca subgen. nov. of Metriogryllacris Karny, 1937. Two genera Gigantogryllacris Karny, 1937 and Pardogryllacris Karny, 1937 are reduced to subgeneric status under Gryllacris Audinet-Serville, 1831 and one monotypic genus Cyanogryllacris Karny, 1937 is synonymised with Capnogryllacris Karny, 1937.        70 species and nine subspecies are described as new: Eugryllacris gandaki sp. nov., Eugryllacris guomashan sp. nov., Eugryllacris crassicauda sp. nov., Eugryllacris crassicauda cambodiana ssp. nov., Eugryllacris serricauda sp. nov., Eugryllacris trabicauda sp. nov., Eugryllacris vermicauda sp. nov., Eugryllacris sulcata sp. nov., Eugryllacris inversa sp. nov., Gryllacris incornuta sp. nov., Gryllacris sok sp. nov., Gryllacris (Pardogryllacris) ovulicauda sp. nov., Gryllacris (Gigantogryllacris) bilineata sp. nov., Gryllacris (Gigantogryllacris) ligulata sp. nov., Lyperogryllacris forcipata sp. nov., Lyperogryllacris ocellata sp. nov., Lyperogryllacris khuntan sp. nov., Ocellarnaca disjuncta sp. nov., Ocellarnaca fusca sp. nov., Otidiogryllacris lawang sp. nov., Otidiogryllacris bamusbama sp. nov., Prosopogryllacris gamta sp. nov., Prosopogryllacris nigra sp. nov., Prosopogryllacris silacea sp. nov., Xanthogryllacris lineata sp. nov., Xanthogryllacris punctata sp. nov., Xanthogryllacris ralum sp. nov., Xanthogryllacris subrecta sp. nov., Aancistroger inarmatus sp. nov., Angustogryllacris bibulbata sp. nov., Aphanogryllacris sinustylata sp. nov., Aphanogryllacris nigritibiae sp. nov., Asarcogryllacris (Asarcogryllacris) brevis sp. nov., Asarcogryllacris (Asarcogryllacris) parapat sp. nov., Asarcogryllacris (Asarcogryllacris) robusta sp. nov., Asarcogryllacris (Pseudolarnaca) cornualis sp. nov., Asarcogryllacris (Pseudasarca) arborea sp. nov., Australogryllacris guttata sp. nov., Capnogryllacris (C.) erythrocephala fuscifrons ssp. nov., Capnogryllacris (C.) varifrons sp. nov., Capnogryllacris (C.) nigromaculata sp. nov., Capnogryllacris (C.) sakaerat toxica ssp. nov., Celebogryllacris brevitegmina sp. nov., Diaphanogryllacris annamita tenera ssp. nov., Diaphanogryllacris opulenta sp. nov., Diaphanogryllacris recta sp. nov., Diaphanogryllacris sinuata sp. nov., Eremus tigris sp. nov., Furcilarnaca chiangdao sp. nov., Furcilarnaca salit sp. nov., Furcilarnaca trilobata sp. nov., Haplogryllacris bilobulata sp. nov., Homogryllacris armigera sp. nov., Homogryllacris stabilis sp. nov., Larnaca (Larnaca) nigricornis sp. nov., Larnaca (Larnaca) tenuis sp. nov., Larnaca (Larnaca) samkos sp. nov., Larnaca (Larnaca) squamiptera sp. nov., Larnaca (Larnaca) subaptera sp. nov., Melaneremus sikkimensis sp. nov., Minigryllacris perpusilla sp. nov., Monseremus appendiculatus sp. nov., Neanias virens sp. nov., Neolarnaca vera nigrinotum ssp. nov., Phryganogryllacris extensa sp. nov., Phryganogryllacris gialaiensis fovealis ssp. nov., Phryganogryllacris nonangulata sp. nov., Siamgryllacris rufa sp. nov., Ultragryllacris pulchra nan ssp. nov., Ultragryllacris triangula sp. nov., Woznessenskia ampliata sp. nov., Woznessenskia bavi sp. nov., Zalarnaca (Glolarnaca) elegantula sp. nov., Zalarnaca (Glolarnaca) globiceps minor ssp. nov., Zalarnaca (Zalarnaca) maninjau sp. nov., Papuogryllacris diluta baiteta ssp. nov., Papuogryllacris rugifrons sp. nov., Papuogryllacris ligata bundi ssp. nov., Paraneanias striatus sp. nov., Plexigryllacris megastyla sp. nov.        The following nomenclatural changes had to be done: Gryllacris bancana Karny, 1930a stat. nov. is raised to full species from subspecies of Gryllacris obscura Brunner von Wattenwyl, 1888; Xanthogryllacris inquinata (Karny, 1928c) comb. stat. nov. is raised to full species from subspecies under Pardogryllacris dyak (Griffini, 1909); Eremus oberthuri Griffini, 1913a stat. nov. is raised to full species from subspecies under Eremus rugosifrons Brunner von Wattenwyl, 1888; Prosopogryllacris nigrovenosa (Karny, 1928b) stat. nov. is raised to full species from subspecies under Prosopogryllacris horvathi (Griffini, 1909); Xanthogryllacris punctipennis aurantiaca (Brunner von Wattenwyl, 1888) stat. reest. and Xanthogryllacris punctipennis dempwolffi (Griffini, 1909b) stat. reest. are treated as valid subspecies under Xanthogryllacris punctipennis (Walker, 1869) instead of being synonyms of the latter; Gryllacris thailandi Gorochov, 2007 and Gryllacris thailandi facemarmiger Gorochov et al., 2015 are downgraded to subspecies of Gryllacris vittata Walker, 1869. Gryllacris kledangensis Karny, 1923 and Gryllacris lombokiana Karny, 1926a become new synonyms of Gryllacris peracca Karny, 1923; Brachyntheisogryllacris abbreviata evolutior (Griffini, 1909a) becomes a new synonym of Brachyntheisogryllacris abbreviata (Brunner von Wattenwyl, 1888), and Haplogryllacris hieroglyphicoides (Chopard, 1924) a new synonym of Haplogryllacris simplex (Walker, 1871).        The following new combinations are proposed: Capnogryllacris (C.) basaliatrata (Griffini, 1909c) comb. nov. from Cyanogryllacris Karny, 1937; Haplogryllacris aliena Walker, 1869 comb. nov. and Haplogryllacris durgensis (Gupta Chandra, 2017) comb. nov. from Gryllacris Audinet-Serville, 1831; Homogryllacris artinii (Griffini, 1913b) comb. nov., Homogryllacris buyssoniana (Griffini, 1912c) comb. nov., Homogryllacris kurseonga (Griffini, 1913b) stat. comb. nov. and Homogryllacris ? maindroni (Griffini, 1913a) comb. nov. from Brachyntheisogryllacris Karny, 1937; Larnaca (Larnaca) eugenii (Griffini, 1914b) comb. nov. from Metriogryllacris Karny, 1937; Larnaca (Larnaca) larnacoides (Karny, 1937) comb. nov. and Larnaca (Larnaca) jacobsoni (Griffini, 1913c) comb. nov. from Melaneremus Karny, 1937.

New taxa and notes on spine-headed katydids (Orthoptera: Conocephalinae: Agraeciini) from the Philippines.

The diversity of the Agraeciini spine-headed katydids (Orthoptera: Tettigonioidea: Conocephalinae) in Southeast Asia remains poorly known, with species still awaiting discovery. Recent field collections in the Philippines resulted in the discovery of three novel species and redescription of two known species of Agraeciini, namely, (1) Anthracites furvuseques sp. nov. from Siargao in Mindanao, a species close to A. major Hebard, 1922; (2) Axylus mabinii sp. nov. from Luzon: (3) a new variant in Axylus c.f. philippinus (Hebard, 1922); (4) Salomona lumadae sp. nov. from Siargao in Mindanao; and (5) Salomona brevicollis Stål, 1877 comb. resurr. is a valid species distinct from Salomona maculifrons Stål, 1877.

A new species of spine-headed katydid Mesagraecia Ingrisch, 1998 (Conocephalinae: Agraeciini), with key to species.

The taxonomy of poorly known Mesagraecia Ingrisch, 1998 is reviewed. A new species of Mesagraecia spine-headed katydid (Conocephalinae: Agraeciini) is described from Bukit Larut, Perak, Peninsular Malaysia: Mesagraecia larutensis sp. n. A key to species is also presented.

A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites Revision of the Indo-Australian Conocephalinae, Part 3.

Axylus group is used to include the five genera Axylus Stål, 1877, Anthracites Redtenbacher, 1891 sensu stricto, Eucoptaspis Willemse, 1966, Eulobaspis gen. nov., and Heminicsara Karny, 1912. It is mainly based on a combination of the characters shape of pronotum, spiniform meso- and metasternal lobes, and similar basic ground plans of the male cerci, titillators and female subgenital plates. The five genera together with two superficially similar genera Euanthracites gen. nov. and Sulasara gen. nov. are fully revised. Papuacites gen. nov. is proposed for two New Guinean species formerly included in Anthracites. Nicsara Walker, 1869 is restricted to Australian species; Spinisternum Willemse, 1942 is synonymised with Heminicsara Karny, 1912; Odontocoryphus Karny, 1907 based on two nymphs is synonymised with Macroxiphus Pictet, 1888; Pseudoliara Karny, 1907 described after one nymph is regarded incertae sedis. 40 new combination of species are proposed: Axylus bimaculatus (Redtenbacher, 1891) comb. nov., A. inferior (Brunner, 1898) comb. nov., A. inflatus (Brunner, 1898) comb. nov., A. loboensis (De Haan, 1842) comb. nov., A. minutus (Dohrn, 1905) comb. nov., A. nigrifrons (Brunner, 1898) comb. nov., A. philippinus (Hebard, 1922) comb. nov., A, taylori (Hebard, 1922) comb. nov., and A. thoracicus (Dohrn, 1905) comb. nov. (all from Nicsara); Euanthracites apoensis (Hebard, 1922) comb. nov., E. femoralis (Dohrn, 1905) comb. nov., E. rufus (Ingrisch, 1998) comb. nov., and E. tibialis (Karny, 1931) comb. nov. (from Anthracites); Eucoptaspis inexpectatus (Willemse, 1953) comb. nov. (from Gonatacanthus Karny, 1907); Eulobaspis dehaani (Karny, 1920) comb. nov., E. emarginata (Karny, 1926) comb. nov., E. moluccana (Redtenbacher, 1891) comb. nov., E. personata (Karny, 1926) comb. nov., E. quadrimaculata (Karny, 1926) comb. nov., E. rotundata (Karny, 1926) comb. nov., and E. strigatipes (Bolivar, 1898) comb. nov. (from Nicsara); Eulobaspis lobaspoides (Karny, 1907) comb. nov. and E. ornata (Brunner, 1898) comb. nov. (from Axylus); Heminicsara excisa (Karny, 1926) comb. nov., H. insulana (Willemse, 1966) comb. nov., H. schlaginhaufeni (Karny, 1912) comb. nov., and H. viridipes (Karny, 1912) comb. nov. (from Nicsara); Heminicsara castaneipictus (Willemse, 1966) comb. nov., H. insularis (Willemse, 1942) comb. nov., and H. palauensis (Vickery & Kevan, 1999) comb. nov. (from Spinisternum); Heminicsara decipiens (Karny, 1926) comb. nov. and H. griffinii (Karny, 1911) comb. nov. (from Gonatacanthus); Heminicsara novaeguineae (Willemse, 1966) comb. nov. (from Eucoptaspis); Sulasara aethiops (Karny, 1931) comb. nov., S. karnyi (Willemse, 1932) comb. nov., and Sulasara sarasini (Karny, 1931) comb. nov. (from Nicsara); Papuacites nigrifrons (Karny, 1912) comb. nov. and P. nakanaiensis (Naskrecki & Rentz, 2010) comb. nov. (from Anthracites); Paramacroxiphus multispinosa (Bolivar, 1898) comb. nov. (from Nicsara); Palaeoagraecia globiceratus Vickery & Kevan, 1999 comb. nov. (from Macroxiphus). Odontocoryphus pullus Karny, 1907 becomes a new synonym of Macroxiphus sumatranus sumatranus (Haan, 1842). 87 species are described as new: nine species in Axylus: A. brachypterus sp. nov., A. dulang sp. nov., A. furcatus sp. nov., A. mengkoka sp. nov., A. montanus sp. nov., A. negros sp. nov. , A. superior sp. nov., A. totop sp. nov. , A. unicolor sp. nov.; six species in Anthracites: A. bilineatus sp. nov., A. flagellatus sp. nov., A. pyramidalis sp. nov., A. romblon sp. nov., A. sinuatus sp. nov., A. unispinus sp. nov.; four species in Euanthracites: E. bispinus sp. nov., E. eboreus sp. nov., E. ile sp. nov., E. uru sp. nov.; six species in Eucoptaspis: E. adonara sp. nov., E. hexamaculatus sp. nov., E. remotus sp. nov., E. stylatus sp. nov., E. trapezoides sp. nov., E. wawo sp. nov.; eight species in Eulobaspis: E. bacan C.Willemse & Ingrisch sp. nov., E. baduri sp. nov., E. buruensis sp. nov., E. ceramica C.Willemse & Ingrisch sp. nov., E. morotai sp. nov., E. sudirman sp. nov., E. ternate sp. nov., E. variata sp. nov.; 51 species in Heminicsara: H. albatros sp. nov., H. albipuncta sp. nov., H. albogeniculata Naskrecki & Ingrisch sp. nov., H. alticola sp. nov., H. ammea sp. nov., H. anggi sp. nov., H. bilobata sp. nov., H. cingima sp. nov., H. comprima sp. nov., H. coriformis sp. nov., H. corneli sp. nov., H. cyclops sp. nov., H. despecta Naskrecki & Ingrisch sp. nov., H. dilatata sp. nov., H. dividata sp. nov., H. dobo sp. nov., H. elongata Naskrecki & Ingrisch sp. nov., H. furcata sp. nov., H. gibba sp. nov., H. gugusu Naskrecki & Ingrisch sp. nov., H. illugi sp. nov., H. jacobii Karny, 1912, H. jayawijaya sp. nov., H. kelila sp. nov., H. kolombangara sp. nov., H. lamas Naskrecki & Ingrisch sp. nov., H. longiloba sp. nov., H. lord sp. nov., H. malu sp. nov., H. mamberamo sp. nov., H. manus sp. nov., H. montana sp. nov., H. nigra sp. nov., H. nomoensis sp. nov., H. obiensis sp. nov., H. ohu sp. nov., H. pak sp. nov., H. parallela Naskrecki & Ingrisch sp. nov., H. pinniger sp. nov., H. popoman sp. nov., H. rugosa sp. nov., H. scutula sp. nov., H. sica sp. nov., H. sinewit sp. nov., H. siwi sp. nov., H. stylata sp. nov., H. tabtab sp. nov., H. truncata Naskrecki & Ingrisch sp. nov., H. tumulus sp. nov., H. umasani sp. nov., H. wanuma sp. nov., H. zugi sp. nov.; and three species in Sulasara: S. armata sp. nov., S. renschi sp. nov., S. tambu sp. nov.

New taxa and notes of some described species of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) from Malay Peninsula.

Four new species from three genera of Agraeciini from Malay Peninsula are described: Paragraecia temasek sp. n., Peracca mirzai sp. n., Peracca macritchiensis sp. n. from Singapore, and Lichnofugia malaya sp. n. from Peninsular Malaysia. The first records and descriptions of the female of Liara alata Ingrisch, 1998 and the male of Paragraecia gracilis Ingrisch, 1998 are given.

New taxa and notes of some described species of scaly crickets (Orthoptera: Mogoplistidae: Mogoplistinae) from Singapore.

Two new species of Mogoplistinae from Singapore are described: Micrornebius kopisua sp. n. and Ornebius insculpta sp. n. The first record and description of males of Ornebius albipalpus Ingrisch, 2006 and females of Ornebius tampines Tan & Robillard, 2012 are given. The occurrence of Ornebius rufonigrus Ingrisch, 1987 in Singapore is confirmed while confirming the identity of another Ornebius species from Singapore with Ornebius cf. pullus Ingrisch, 2006 requires further investigation. A key to the species of Mogoplistinae from Singapore is provided.

The plurennial life cycles of the European Tettigoniidae (Insecta: Orthoptera) : 1. The effect of temperature on embryonic development and hatching.

The effect of temperature on embryonic development, voltinism, and hatching was studied in the laboratory in eggs of 21 Central and Southeastern European Tettigoniidae species. In most species, the embryo has to arrive at a postkatatrepsis stage prior to the onset of cold to be able to hatch in the following spring. The rate of embryonic development differs: quickly developing species need 4 weeks at 24°C (prior to cold) and almost all eggs hatch after the first cold treatment, slowly developing species would need 8-12 weeks to do the same. In Central Europe, warmth is not enough for the slowly developing species to have an univoltine life cycle, but they could have it in southern Europe. Most species make use of a dormancy sequence to pass successive winters as follows: an initial embryonic dormancy (either quiscence or diapause in embryonic stage 4) and a final diapause in embryonic stage 23/24. Additionally, 3 forms of aestivation or summer dormancy were observed facultatively: an initial diapause in embryonic stage 4 (induced and terminated at 30°C), a median dormancy shortly before or after katatrepsis (at 30°C), and a penultimate diapause in embryonic stage 20 (at 24°C).The life cycles of the European Tettigoniidae species can follow one of 3 types: 1. annual life cycle (no initial embryonic dormancy); 2. annual or biennial depending on whether laid early or late; 3. biennial or many year life cycle (up to 8 years due to a prolonged initial diapause).

The plurennial life cycles of the European Tettigoniidae (Insecta: Orthoptera) : 2. The effect of photoperiod on the induction of an initial diapause.

In eggs of European Tettigoniidae species, an initial diapause can occur just after blastoderm formation and a final diapause close to the end of embryonic development. The effect of photoperiod experienced maternally on the induction of the initial diapause was studied, using 13 species from Central and Southeastern Europe. In Conocephalus dorsalis and Platycleis albopunctata there was no initial diapause induced by photoperod, while, in Tettigonia-and Saga-species and in Metrioptera saussuriana it occurred independently of day length. In Metrioptera roeseli, M. bicolor, Leptophyes punctatissima, Eupholidoptera smyrnensis, and Decticus verrucivorus, oviposition at short day induced an initial diapause, while after oviposition at long day, the eggs developed directly until final diapause. On the other hand, in up to 1/4 of the eggs of Pholidoptera griseoaptera an initial diapause was induced by long day length. Populations of D. verrucivorus from different latitudes differed with respect to the critical day length. For E. smyrnensis and M. roeseli, a photoperiodic response curve was calculated.In the Rhodian population of E. smyrnensis, the dormancy sequence of initial and final embryonic diapause can be used for aestivation and hibernation within an annual life cycle, while enabling hibernation in successive years for the Central European species. The population of D. verrucivorus near Aachen has an "obligatory" plurennial life cycle, since the critical day length for development without initial diapause is above the range of day length occurring in the field.

The plurennial life cycles of the European Tettigoniidae (Insecta: Orthoptera) : 3. The effect of drought and the variable duration of the initial diapause.

The effect of drought on embryonic development and on hatching was studied in 13 European Tettigoniidae species. Drought can affect development in three different ways: (1) Embryonic development proceeds slower than if the eggs are in contact with water; (2) it stops (for final diapause) in an earlier embryonic stage; (3) it affects maintenance and termination of the initial embryonic diapause.In many Tettigoniidae species, the initial diapause is prolonged, and may last several years. Without draught stress, between 1 and 7 cold treatments in the laboratory, and with eggs of the Tettigonia-species between 1 and 6 winters in the field were necessary to enable all eggs to complete initial diapause. In Central European species, the number of eggs maintaining initial diapause significantly increased when the eggs had no contact with water at the time when they should recover from diapause. In contrast, termination of initial diapause in Tettigonia caudata from Greece, when the environment became favorable for growth again, was highest in that group of eggs that had lost most water in a preceding period of drought. The importance of the prolonged initial diapause for the survival of unpredictable adverse conditions is discussed.

Effect of hibernation length on termination of diapause in European Tettigoniidae (Insecta: Orthoptera).

The effect of the length of cold treatment on development time of embryos of eight Central European tettigoniid species was investigated in laboratory experiments, using chilling periods of 3, 4, 5, 6, 9, and 12 months. Two different types of response were observed: (1) The hatching date was not influenced by hibernation length (Conocephalus) or only after the 3-month cold treatment (Phaneroptera, Platycleis), or (2) The time required for post-diapause development was significantly reduced by each 1-month increase in the length of cold treatment up to 6 months (Meconema, Metrioptera, Tettigonia).Thus, the correct timing of hatching is achieved by one of two different strategies. In lat-hatching species, the influence of hibernation length is small. Premature appearance is prevented by a high thermal threshold. In early-hatching species, the embryos are obviously able to measure the length of the chilling period. Thus, they gradually change their physiological state so that the potential speed of postdiapause development increases as the 'winter' is prolonged. This strategy can be understood as an adaptation to the necessity of hatching at low or moderate temperatures. Field experiments with two Tettigonia species confirm the laboratory data.

The influence of environmental factors on dormancy and duration of egg development in Metrioptera roeseli (Orthoptera: Tettigoniidae).

Developing eggs of Metrioptera roeseli HGB. have an obligate "embryonic" diapause in stage 23 and a facultative "initial" diapause in stage 4. Initial diapause is lacking when the ovipositing females are kept at long day photoperiod (LD 16/8), but is induced at LD 14/10 and LD 8/16. When natural light conditions are given in the laboratory, the females mostly lay diapausing eggs from the end of August onwards. In eggs without initial diapause, a larger number of embryos reach a stage, in which the embryonic diapause can be broken by chilling, only at favorable temperature (4 weeks at 24°C or 8 at 18°C) and in contact with water before the first winter. Thus, some individuals of a given population of M. roeseli have an annual, others a biennial life cycle. The variability in the duration of egg development is due to a photoperiodically induced, facultative initial diapause and to a possible quiescence caused by insufficient temperature and/or moisture.